Alyssa Milano Is the One Who Got Away on 'Castle'

Despite his playboy ways, Richard Castle (Nathan Fillion) was once in love. Check out the video to see Alyssa Milano as the one who got away!

When Castle and Beckett (Stana Katic) are called in to investigate the death of a bridesmaid at a posh wedding, Castle is stunned to discover that the soon-to-be bride is his ex, Kyra (Milano).

"Of all of the things you worry about going wrong at your wedding…," Kyra tells Castle.